Untuned organ mutation stops generate audible beats when chords are played

Evidence

For example, if a C is played with 8′ and 2/3′ stops drawn, the f0 of the C on the 22/3′ rank will be exactly in tune with the third harmonic of the C on the 8′ rank. If the G above is now played also to produce a two-note chord, the first harmonic of the G on the 8′ rank will beat with the f0 of the C on the 22/3′ rank (and the third harmonic of the C on the 8′ rank). — David M. Howard, Jamie Angus, Acoustics and Psychoacoustics, 1561

Categorization

• QUESTION: Could trace exactly which harmonic combinations produce which beating rates to build a mental model of organ stop interactions.